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A book lung is a type of respiration organ used for atmospheric gas exchange present in some arachnids such as scorpions and tetrapulmonates. It is a terrestrial adaptation of the book gills seen in aquatic chelicerates such as horseshoe crabs, where the gills are ventral appendages under the abdomen (opisthosoma) arranged into a series of page-like lamellae in order to maximize surface area for diffusion. In arachnids, each of these organs is housed inside an air-filled cavity (atrium) that opens to the surrounding atmosphere through a small opening (spiracle), which provides a controlled environment for respiration.
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