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In zoology, copulation is animal sexual behavior in which a male introduces sperm into the female's body, especially directly into the female's reproductive tract. This is an aspect of mating. Many aquatic animals use external fertilization, whereas internal fertilization may have developed from a need to maintain gametes in a liquid medium in the Late Ordovician epoch. Internal fertilization with many vertebrates occurs via cloacal copulation, known as cloacal kiss (see also hemipenis), while most mammals copulate vaginally, and many basal vertebrates reproduce sexually with external fertilization.
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