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Soil harbours a huge number of animal species, whether over their entire life or at least during larval stages. Soil offers protection against environmental hazards, such as excess temperature and moisture fluctuations, in particular in arid and cold environments, as well as against predation. Soil provisions food over the year, especially since omnivory seems the rule rather than the exception, and allows reproduction and egg deposition in a safe environment, even for those animals not currently living belowground. Many soil invertebrates, and also some soil vertebrates, are tightly adapted to a subterranean concealed environment, being smaller, blind, depigmented, legfree or with reduced legs, and reproducing asexually, with negative consequences on their colonization rate when the environment is changing at landscape scale. It has been argued that soil could have been a crucible for the evolution of invertebrate terrestrial faunas, as an intermediary step in the transition from aquatic to aerial life.
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